A revision of the new Andean butterfly genus Optimandes Marín, Nakahara & Willmott, n. gen., with the description of a new species (Nymphalidae: Satyrinae: Euptychiina)


  • Keith R. Willmott McGuire Center for Lepidoptera and Biodiversity Florida Museum of Natural History
  • Mario A. Marín
  • Shinichi Nakahara
  • Tatiana Pomerantz
  • Gerardo Lamas
  • Blanca Huertas
  • Marianne Espeland
  • Lei Xiao
  • Jason P. W. Hall
  • James I. Robinson Willmott
  • André V. L. Freitas


Andes, Chusquea, immature stages, inventory, mimicry, species description, taxonomy


A new genus, Optimandes Marín, Nakahara & Willmott, n. gen., is described to contain the type species Neonympha eugenia C. Felder & R. Felder, 1867, its junior subjective synonym Euptychia phineus Butler, 1867, and its subspecies Euptychia transversa Weymer, 1911, which are transferred from the genus Euptychoides Forster, 1964. The sister species to Optimandes eugenia n. comb. is a distinctive new species which is here described and named as Optimandes mocha Willmott, Hall & Lamas, n. sp.. Figures of wings, genitalia and distribution maps are provided for both species and the immature stages of Optimandes eugenia are described for the first time. Both species are uncommon to rare inhabitants of Andean cloud forest, with O. eugenia widespread throughout the tropical Andes, and O. mocha occurring from southern Ecuador to Bolivia.