Spatial Patterns of Canopy Leaves of Saplings of Four Woody Species in a Forest Environment


forest environment
spatial pattern

How to Cite

Li, D., & Wu, S. (2000). Spatial Patterns of Canopy Leaves of Saplings of Four Woody Species in a Forest Environment. Selbyana, 21(1/2), 88–96. Retrieved from


The spatial patterns of canopy leaves of saplings of four woody species in a natural secondary forest of northeastern China were analyzed. Four saplings displayed leaf area in higher layers with larger positive inclination angles more than that of the other layers. However, this proportion declined gradually along with descending sapling height, and the leaf area with smaller inclination angles gradually increased. Along the gradient of azimuth angle, the frequency distribution patterns of the leaves of four saplings were irregular and stochastic to some extent. The leaf area density curves of two saplings were parabola-shaped along the gradient of sapling height, and in the middle layers the leaf area densities were higher than those of lower and top layers in sequence. Applying the formulae of Levins (1968) and Shannon and Weaver (1949), as well as the formulae of Schoener (1970) and Pianka (1974), the characteristics of leaf populations of four saplings distributed along the gradients of inclination and azimuth angles, were further analyzed. The significance of these adaptive characteristics in understanding the dynamic properties and regeneration process of the forest community are discussed.


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